Bee (trigona minima)

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Bees are flying insects, closely related to wasps and ants. Bees are a monophyletic lineage within the superfamily Apoidea, presently classified by the unranked taxon name Anthophila. There are slightly fewer than 20,000 known species of bees, though many are undescribed and the actual number is probably higher. They are found on every continent except Antarctica.
Many species of bees are poorly known. The smallest bee is the dwarf bee (Trigona minima) and it is about 2.1 mm (5/64") long. The largest bee in the world is Megachile pluto, which can be as large as 39 mm (1.5"). The most common type of bee in the Northern Hemisphere are the many species of Halictidae, or sweat bees, though this may come as a surprise to people, as they are small and often mistaken for wasps or flies. The most well-known bee species is the Western honey bee, which, as its name suggests, produces honey, as do a few other types of bee. Human control of this species is known as beekeeping or apiculture.
Bees are adapted for feeding on nectar and pollen, the former primarily as an energy source, and the latter primarily for protein and other nutrients. Most pollen is used as food for larvae.

Bees have a long proboscis that enables them to obtain the nectar from flowers. Bees have antennae almost universally made up of thirteen segments in males and twelve in females, as is typical for the superfamily. They all have two pairs of wings, the back pair being the smaller of the two; in a very few species, one sex or caste has relatively short wings that make flight difficult or impossible, but none are wingless.

Pollination
Bees play an important role in pollinating flowering plants, and are the major type of pollinators in ecosystems that contain flowering plants. Bees may focus on gathering nectar or on gathering pollen, depending on their greater need at the time. Bees gathering nectar may accomplish pollination, but bees that are deliberately gathering pollen are more efficient pollinators. It is estimated that one third of the human food supply depends on insect pollination, most of this accomplished by bees.

Most bees are fuzzy and carry an electrostatic charge, thus aiding in the adherence of pollen. Female bees periodically stop foraging and groom themselves to pack the pollen into the scopa, which is on the legs in most bees, and on the ventral abdomen on others, and modified into specialized pollen baskets on the legs of honey bees and their relatives. Many bees are opportunistic foragers, and will gather pollen from a variety of plants, but many others are oligolectic, gathering pollen from only one or a few types of plant. A small number of plants produce nutritious floral oils rather than pollen, which are gathered and used by oligolectic bees. One small subgroup of stingless bees (called "vulture bees") is specialized to feed on carrion, and these are the only bees that do not use plant products as food. Pollen and nectar are usually combined together to form a "provision mass", which is often soupy, but can be firm. It is formed into various shapes (typically spheroid), and stored in a small chamber (a "cell"), with the egg deposited on the mass. The cell is typically sealed after the egg is laid, and the adult and larva never interact directly (a system called "mass provisioning").

Bees are extremely important as pollinators in agriculture, especially the domesticated Western honey bee, with contract pollination having overtaken the role of honey production for beekeepers in many countries. Monoculture and pollinator decline have increasingly caused honey bee keepers to become migratory so that bees can be concentrated in areas of pollination need at the appropriate season. Many other species of bees are increasingly cultured and used to meet the agricultural pollination need. Bees also play a major, though not always understood, role in providing food for birds and wildlife. Many of these bees survive in refuge in wild areas away from agricultural spraying, only to be poisoned in massive spray programs for mosquitoes, gypsy moths, or other pest insects.

Visiting flowers is a dangerous occupation with high mortality rates. Many assassin bugs and crab spiders hide in flowers to capture unwary bees. Others are lost to birds in flight. Insecticides used on blooming plants can kill large numbers of bees, both by direct poisoning and by contamination of their food supply. A honey bee queen may lay 2000 eggs per day during spring buildup, but she also must lay 1000 to 1500 eggs per day during the foraging season, simply to replace daily casualties. The population value of bees depends partly on the individual efficiency of the bees, but also on the population itself. Thus, while bumblebees have been found to be about ten times more efficient pollinators on cucurbits, the total efficiency of a colony of honey bees is much greater, due to greater numbers. Likewise, during early spring orchard blossoms, bumblebee populations are limited to only a few queens, thus they are not significant pollinators of early fruit.

Evolution
Bees, like ants, are essentially a highly specialized form of wasp. The ancestors of bees were wasps in the family Crabronidae, and therefore predators of other insects. The switch from insect prey to pollen may have resulted from the consumption of prey insects that were flower visitors and were partially covered with pollen when they were fed to the wasp larvae. This same evolutionary scenario has also occurred within the vespoid wasps, where the group known as "pollen wasps" also evolved from predatory ancestors. The oldest bee fossil, of the genus Melittosphex, is 100 million years old and supports the theory that bees evolved from wasps, and subsequently evolved alongside flowers. The earliest animal pollinated flowers were pollinated by insects such as beetles, so the syndrome of insect pollination was well established before bees first appeared. The novelty is that bees are specialized as pollination agents, with behavioral and physical modifications that specifically enhance pollination, and are much more efficient at the task than beetles, flies, butterflies, pollen wasps, or any other pollinating insect. The appearance of such floral specialists is believed to have driven the adaptive radiation of the angiosperms, and, in turn, the bees themselves.